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Aging and Gerontology

Cell and Molecular Biology Graduate Program

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Ecology, Evolution, and Conservation Biology

Marine Biology

Neurosciences Graduate Specialization

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International Cultural Studies

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Resource Management


Marine Biology

c/o Hawai'i Institute of Marine Biology (HIMB)
P.O. Box 1346
Kaneohe, HI 96744
Tel: (808) 236-7401
Fax: (808) 236-7443
Web: www.hawaii.edu/marine_biology/graduate

Graduate Faculty

J. Leong, PhD (Chair)—HIMB
M. Alam, PhD—microbiology
W. W. L. Au, PhD—HIMB
J. H. Bailey-Brock, PhD—zoology
R. Bidigare, PhD—HIMB
C. Birkeland, PhD—zoology
B. Bowen, PhD—HIMB
R. Cann, PhD—genetics
S. Conant, PhD—zoology
I. Cooke, PhD—zoology
H. G. de Couet, PhD—zoology
S. Donachie, PhD-microbiology
J. Douglas, PhD—microbiology
R. Gates, PhD—HIMB
G. Grau, PhD—zoology
K. Holland, PhD—HIMB
T. D. Humphreys, PhD—biochemistry
C. Hunter, PhD—biology
D. Jameson, PhD—biochemistry
P. Jokiel, PhD—HIMB/zoology
D. Karl, PhD—oceanography
S. Karl, PhD—HIMB
R. Kinzie III, PhD—zoology
P. E. Nachtigall, PhD—HIMB
M. Rappe, PhD—HIMB
K. Selph, PhD—oceanography
C. Smith, PhD—botany
G. Steward, PhD—biological oceanography
F. Thomas, PhD—HIMB
R. Toonen, PhD—HIMB
T. Tricas, PhD—zoology
G. Wang, PhD—oceanography

The Academic Program

The marine biology specialization is a UH Manoa-wide program focusing on recent advances in the understanding of marine systems at the ecological, organismic, and cellular-molecular levels. Students can select courses, advisors, and research opportunities from a wide range of specialties, including: marine botany, ecology, genetics, virology, microbiology, zoology, aquaculture, behavioral biosystematics, biological oceanography, coral reef biology, fisheries, and molecular biology.

The marine biology specialization is available to graduate students in botany, microbiology, oceanography, and zoology. Prospective graduate students must apply first to one of these programs. Applications from students who have been accepted to botany, microbiology, oceanography, or zoology who have also indicated a desire to specialize in marine biology are reviewed by the Marine Biology Admissions Committee.

Students specializing in marine biology supplement the courses required for a degree in their chosen field with courses specific to marine biology. The actual selection is determined by the student in consultation with his or her advisor. Graduate student research is carried out in the laboratories of the graduate faculty. These include laboratories in Edmondson Hall, Snyder Hall, St. John Laboratory (botanical sciences), Marine Science Building, Hawai'i Institute of Marine Biology (located on Coconut Island in Kaneohe Bay), Bekesy Laboratory, and Kewalo Laboratory of the Pacific Biomedical Research Center. Research capabilities include DNA sequencing using PCR technology; video and acoustic recording for ecological and behavioral studied of coral reef and planktonic organisms; transmission and scanning electron, ultraviolet, and light microscopy; electrophoretic analysis; flow cytometry; and radioisotope tracer studies.

Selected courses:

  • ANSC 450 Aquaculture Production (3)
  • BOT 480 Algal Diversity and Evolution (4)
  • BOT 680 Marine Macrophytes Seminar (2)
  • OCN 450 Aquaculture Production (3)
  • OCN 621 Biological Oceanography (3)
  • OCN 626 Marine Microplankton Ecology (4)
  • OCN 627 Ecology of Pelagic Marine Animals (4)
  • OCN 628 Benthic Biological Oceanography (4)
  • OCN 750 Topic in Biological Oceanography (V)
  • ZOOL 466 Fisheries Science (3)
  • ZOOL 467 Ecology of Fishes (3)
  • ZOOL 475 Biology of the Invertebrates (3)
  • ZOOL 620 Marine Ecology (3)

Neurosciences Graduate Specialization

Graduate Faculty

M. Berry, PhD (Chair)—cell and molecular biology
D. C. Blanchard, PhD—cell and molecular biology
R. Blanchard, PhD—psychology
T. Blank, PhD—cell and molecular biology
L. Chang, MD—medicine
A. Dunn, PhD—Bekesy Laboratory of Neurobiology
T. Ernst, PhD—medicine
A. Fleig, PhD—cell and molecular biology
M. Gerschenson, PhD—medicine
D. Hartline, PhD—Bekesy Laboratory of Neurobiology
V. Nerurkar, PhD—tropical medicine and medical microbiology
R. Nichols,* PhD—cell and molecular biology
H. Petrovitch, MD—geriatric medicine
M. Rayner,* PhD—cell and molecular biology
G. Webster Ross, MD—medicine and geriatric medicine
C. Shikuma, MD—medicine
B. Shiramizu, MD—medicine/pediatrics
A. Stenger, PhD—medicine
L. Takahashi, PhD—psychology
L. White, MD., PhD—pediatrics


The purpose of this specialization is to allow graduate students pursuing masters and doctoral degrees in various disciplines (e.g., medicine, psychology, engineering, computer science, molecular bioscience, etc.) to formally specialize in neuroscience. This is accomplished by taking a minimum of 4 courses plus participate in a "journal club" focusing on neuroscience as approved by the Graduate Chair.


  • CMB 606 Introduction to Neurosciences (4)
  • CMB 621 Cell Molecular Biology I (4)
  • CMB 622 Cell Molecular Biology II (4)
  • CMB 626 Ethics in Biomedical Research (2)
  • CMB 705 Special Topics in Neurosciences (V)
  • PHRM 640 Neuropharmacology (2
  • PSY 622 Animal Learning (3)
  • PSY 626 Cognitive Psychology (3)
  • PSY 631 Comparative Psychology (3)
  • PSY 642 Cognitive Development (3)
  • PSY 719 Research in Psychometrics (3)
  • TRMD 607 Neurovirology (1)
  • ZOOL 642 Cellular Neurophysiology (3)
  • ZOOL 712 Topics in Nerve/Muscle Physiology (V)

CMB Courses

Interdisciplinary Studies

Degree Offered: BA in interdisciplinary studies

See the “Colleges of Arts and Sciences” section for more information.

IS Courses